Redigerer Orphnus

{{Taksoboks
| navn            =''Orphnus''
| bilde           =
| bildetekst      =
| norsknavn       =
| vitenskapsnavn  =''Orphnus''
| autor           = W.S. MacLeay
| autorår         = 1819
| regnum = [[Dyreriket]]
| phylum = [[Leddyr]]
| classis = [[Insekter]]
| ordo = [[Biller]]
| familia = [[skarabider]] (Scarabaeidae)
| subfamilia = [[Orphninae]]
}}
{{Taksoboks-gruppe
| artstall        = over 100
| habitat         =
| utbredelse      =Afrika}}
* Se systematisk inndeling
{{Taksoboks-slutt}}

'''''Orphnus''''' er en artsrik slekt av [[biller]] som hører til familien [[skarabider]] (Scarabaeidae).

== Utseende ==
Små til ganske store (5-25 millimeter), bredt ovale, brune til svarte [[skarabider]]. Antennene er 10-leddete. Pronotum kan ha opphøyde knøler eller være uthulet i midten, hannene hos noen arter har en knøl eller et horn i pannen. Beian er kraftige med brede tenner på for-leggene og lange sporer på bakleggene.

== Levevis ==
Lite er kjent om biologien til disse billene, som har et mer eller mindre underjordisk levesett. De voksne billene er ikke flygedyktige.

== Utbredelse ==
De aller fleste av artene lever i Afrika, men det finnes også seks arter i det sørlige og sørøstlige Asia.

== Systematisk inndeling ==

{{tre}}
* Ordenen [[biller]], Coleoptera
** Underordenen [[Polyphaga]]
*** Overfamilien [[Skarabider (Scarabaeoidea)|skarabider]], Scarabaeoidea
**** Familien [[skarabider]], (Scarabaeidae) <small>[[Latreille]], 1806</small>
***** Underfamilie [[Orphninae]] <small>Erichson, 1847</small>
****** Stammen [[Orphnini]]
******* Slekten '''''Orphnus''''' <small>MacLeay, 1819</small> 
******** Underslekten ''[[Cerhomalus]]'' <small>Quedenfeldt, 1884</small> - noen ganger regnet som en egen slekt
********* ''[[Orphnus (Cerhomalus) absconditus]]'' <small>(Petrovitz, 1971)</small>
********* ''[[Orphnus (Cerhomalus) mechowi]]'' <small>(Quedenfeldt, 1884)</small>
******** Underslekten ''[[Horpnus]]'' <small>Paulian, 1948</small>
********* ''[[Orphnus (Horpnus) asperatus]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus (Horpnus) ellenbergeri]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus (Horpnus) zumpti]]'' <small>Petrovitz, 1963</small>
******** Underslekten ''Orphnus'' (i snever forstand)
********* ''[[Orphnus acuticornis]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus amplitarsis]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus angolensis]]'' <small>Quedenfeldt, 1884</small>
********* ''[[Orphnus babaulti]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus benderitteri]]'' <small>Pic, 1930</small>
********* ''[[Orphnus bicolor]]'' <small>([[Fabricius]], 1801)</small>
********* ''[[Orphnus bifidus]]'' <small>Schmidt, 1912</small>
********* ''[[Orphnus bilobus]]'' <small>Klug, 1855</small>
********* ''[[Orphnus brunneus]]'' <small>Benderitter, 1912</small>
********* ''[[Orphnus camerunensis]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus capensis]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus chappuisi]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus compactilis]]'' <small>Quedenfeldt, 1884</small>
********* ''[[Orphnus compactus]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus compressicornis]]'' <small>Benderitter, 1913</small>
********* ''[[Orphnus congolanus]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus convexus]]'' <small>Benderitter, 1913</small>
********* ''[[Orphnus coprodoides]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus costatus]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus crassus]]'' <small>Pic, 1928</small>
********* ''[[Orphnus cribratellus]]'' <small>Fairmaire, 1878</small>
********* ''[[Orphnus declivis]]'' <small>Schmidt, 1912</small>
********* ''[[Orphnus drumonti]]'' <small>Frolov, 2009</small>
********* ''[[Orphnus fossatus]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus foveolatus]]'' <small>Benderitter, 1920</small>
********* ''[[Orphnus galla]]'' <small>Gestro, 1895</small>
********* ''[[Orphnus gilleti]]'' <small>Benderitter, 1923</small>
********* ''[[Orphnus grossepunctatus]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus guineensis]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus harrisoni]]'' <small>Frolov, 2008</small>
********* ''[[Orphnus herero]]'' <small>Petrovitz, 1963</small>
********* ''[[Orphnus heteronychoides]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus imitator]]'' <small>Benderitter, 1920</small>
********* ''[[Orphnus impressus]]'' <small>Westwood, 1845</small>
********* ''[[Orphnus incultus]]'' <small>(Péringuey, 1892)</small>
********* ''[[Orphnus jeanneli]]'' <small>Benderitter, 1914</small>
********* ''[[Orphnus kafuensis]]'' <small>Péringuey, 1908</small>
********* ''[[Orphnus leleupi]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus letestui]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus longicornis]]'' <small>Frolov & Akhmetova, 2015</small>
********* ''[[Orphnus luluanus]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus luminosus]]'' <small>Benderitter, 1920</small>
********* ''[[Orphnus macleayi]]'' <small>Laporte de Castelnau, 1832</small>
********* ''[[Orphnus mandibularis]]'' <small>(Lansberge, 1886)</small>
********* ''[[Orphnus mashunensis]]'' <small>Péringuey, 1908</small>
********* ''[[Orphnus mombasaensis]]'' <small>Benderitter, 1914</small>
********* ''[[Orphnus mpese]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus mysoriensis]]'' <small>Westwood, 1845</small>
********* ''[[Orphnus niger]]'' <small>Pic, 1928</small>
********* ''[[Orphnus nyassicus]]'' <small>Kolbe, 1895</small>
********* ''[[Orphnus orbus]]'' <small>Benderitter, 1920</small>
********* ''[[Orphnus oryctoides]]'' <small>Quedenfeldt, 1888</small>
********* ''[[Orphnus ovampoanus]]'' <small>Péringuey, 1896</small>
********* ''[[Orphnus overlaeti]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus parentalis]]'' <small>Péringuey, 1908</small>
********* ''[[Orphnus parvus]]'' <small>Wiedemann, 1823</small>
********* ''[[Orphnus pauliani]]'' <small>Gomes Alves, 1957</small>
********* ''[[Orphnus pici]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus picinus]]'' <small>Westwood, 1845</small>
********* ''[[Orphnus pugnax]]'' <small>Péringuey, 1896</small>
********* ''[[Orphnus rufithorax]]'' <small>Benderitter, 1914</small>
********* ''[[Orphnus rufulus]]'' <small>Boheman, 1857</small>
********* ''[[Orphnus sansibaricus]]'' <small>Kolbe, 1895</small>
********* ''[[Orphnus schoutedeni]]'' <small>Benderitter, 1920</small>
********* ''[[Orphnus senegalensis]]'' <small>Laporte de Castelnau, 1832</small>
********* ''[[Orphnus similis]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus sinuaticeps]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus sinuatus]]'' <small>Benderitter, 1923</small>
********* ''[[Orphnus striatoides]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus striatopunctatus]]'' <small>Felsche, 1904</small>
********* ''[[Orphnus striatus]]'' <small>Benderitter, 1913</small>
********* ''[[Orphnus subcornutus]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus subfurcatus]]'' <small>Kolbe, 1895</small>
********* ''[[Orphnus sudanicus]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus tchadensis]]'' <small>Benderitter, 1913</small>
********* ''[[Orphnus thoracicus]]'' <small>Linell, 1896</small>
********* ''[[Orphnus tinantae]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus transvaalensis]]'' <small>Frolov, 2009</small>
********* ''[[Orphnus tristis]]'' <small>Pic, 1928</small>
********* ''[[Orphnus usambaricus]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus viduae]]'' <small>Petrovitz, 1971</small>
******** Underslekten ''[[Pachyorphnus'']] <small>Paulian, 1948</small>
********* ''[[Orphnus (Pachyorphnus) clavipes]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus (Pachyorphnus) dewittei]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus (Pachyorphnus) subfoveatus]]'' <small>Fairmaire, 1898</small>
******** Underslekten ''[[Phornus]]'' <small>Paulian, 1948</small> <ref>Frolov, A.V. & Akhmetova, L.A. (2016) Revision of the subgenus ''Orphnus'' (''Phornus'') (Coleoptera, Scarabaeidae, Orphninae). ''European Journal of Taxonomy'' 241: 1–20</ref> 
********* ''[[Orphnus ferrierei]]'' <small>Frolov & Akhmetova, 2016</small>
********* ''[[Orphnus giganteus]]'' <small>Paulian, 1948</small>
********* ''[[Orphnus parastrangulatus]]'' <small>Frolov & Akhmetova, 2016</small>
********* ''[[Orphnus renaudi]]'' <small>Frolov & Akhmetova, 2016</small>
********* ''[[Orphnus strangulatus]]'' <small>Paulian, 1948</small>
********* ''[[Orphnus valeriae]]'' <small>Frolov & Akhmetova, 2016</small>
******** Underslekten ''[[Ronphus]]'' <small>Paulian, 1948</small>
********* ''[[Orphnus (Ronphus) demeyeri]]'' <small>Frolov, 2013</small>
********* ''[[Orphnus (Ronphus) livingstonei]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus (Ronphus) massarti]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus (Ronphus) peringueyi]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus (Ronphus) planicollis]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus (Ronphus) quadrigibbosus]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus (Ronphus) simonii]]'' <small>Petrovitz, 1971</small>
********* ''[[Orphnus (Ronphus) testaceus]]'' <small>Paulian, 1951</small>
******** ikke plassert til underslekt
********* ''[[Orphnus audeoudi]]'' <small>Petrovitz, 1975</small>
********* ''[[Orphnus plebejus]]'' <small>Péringuey, 1901</small>
********* ''[[Orphnus posthi]]'' <small>Paulian, 1951</small>
********* ''[[Orphnus rufus]]'' <small>Guérin-Ménéville, 1844</small>

== Referanser ==
<references/>

== Kilder ==
* [http://www.scarabaeoidea.com/taxa/orphninae/taxa-pages/orphnus Nettside om slekten] {{Wayback|url=http://www.scarabaeoidea.com/taxa/orphninae/taxa-pages/orphnus |date=20170208033543 }}
* [http://insecta.pro/taxonomy/187393 Oversikt over artene]
* [http://www.scarabaeoidea.com/taxa/orphninae/catalogue Katalog over artene] {{Wayback|url=http://www.scarabaeoidea.com/taxa/orphninae/catalogue |date=20170211080016 }}

== Eksterne lenker ==
* {{Artslenker}}
* {{Commonscat|Orphninae}}

{{Autoritetsdata}}
{{Kursiv tittel}}

[[Kategori:Orphninae]]
[[Kategori:Biller formelt beskrevet i 1819]]
[[Kategori:Dyr formelt beskrevet av William Sharp Macleay]]